/usr/share/pyshared/brian/stdp.py is in python-brian 1.3.1-1build1.
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Spike-timing-dependent plasticity
'''
# See BEP-2-STDP
from inspection import *
from equations import *
from monitor import SpikeMonitor, RecentStateMonitor
from network import NetworkOperation
from neurongroup import NeuronGroup
from stateupdater import get_linear_equations, LinearStateUpdater
from scipy.linalg import expm
from scipy import dot, eye, zeros, array, clip, exp, Inf
from stdunits import ms
from connections import DelayConnection, DenseConstructionMatrix, SparseConnectionVector
import re
from utils.documentation import flattened_docstring
from copy import copy
import warnings
from itertools import izip
from numpy import arange, floor
from clock import Clock
from units import second
from utils.separate_equations import separate_equations
from log import *
from globalprefs import *
from optimiser import freeze
__all__ = ['STDP', 'ExponentialSTDP']
class STDPUpdater(SpikeMonitor):
'''
Updates STDP variables at spike times
'''
def __init__(self, source, C, vars, code, namespace, delay=0 * ms):
'''
source = source group
C = connection
vars = variable names
M = matrix of the linear differential system
code = code to execute for every spike
namespace = namespace for the code
delay = transmission delay
'''
super(STDPUpdater, self).__init__(source, record=False, delay=delay)
self._code = code # update code
self._namespace = namespace # code namespace
self.C = C
def propagate(self, spikes):
if len(spikes):
self._namespace['spikes'] = spikes
self._namespace['w'] = self.C.W
exec self._code in self._namespace
class DelayedSTDPUpdater(SpikeMonitor):
def __init__(self, C, reverse, delay_expr, max_delay,
vars, other_vars, varmon, othervarmon, code, namespace, delay=0 * ms):
if reverse:
source = C.target
self.get_times_seq = 'get_cols'
else:
source = C.source
self.get_times_seq = 'get_rows'
super(DelayedSTDPUpdater, self).__init__(source, record=False, delay=delay)
self._code = code # update code
self._namespace = namespace # code namespace
self.C = C
self.vars = vars
self.other_vars = other_vars
self.varmon = varmon
self.othervarmon = othervarmon
delay_expr = re.sub(r'\bmax_delay\b', str(float(max_delay)), delay_expr)
delay_expr = 'lambda d:' + delay_expr
self.delay_expr = eval(delay_expr)
def propagate(self, spikes):
if len(spikes):
if isinstance(self.get_times_seq, str):
self.get_times_seq = getattr(self.C.delayvec, self.get_times_seq)
times_seq = self.get_times_seq(spikes)
times_seq = [self.delay_expr(times) for times in times_seq]
for var in self.other_vars:
delayed_values = self.othervarmon[var].get_past_values_sequence(times_seq)
self._namespace[var + '__delayed_values_seq'] = delayed_values
self._namespace['spikes'] = spikes
self._namespace['w'] = self.C.W
exec self._code in self._namespace
class STDP(NetworkOperation):
'''
Spike-timing-dependent plasticity
Initialised with arguments:
``C``
Connection object to apply STDP to.
``eqs``
Differential equations (with units)
``pre``
Python code for presynaptic spikes, use the reserved symbol ``w`` to
refer to the synaptic weight.
``post``
Python code for postsynaptic spikes, use the reserved symbol ``w`` to
refer to the synaptic weight.
``wmin``
Minimum weight (default 0), weights are restricted to be within this
value and wmax.
``wmax``
Maximum weight (default unlimited), weights are restricted to be within
wmin and this value.
``delay_pre``
Presynaptic delay
``delay_post``
Postsynaptic delay (backward propagating spike)
The STDP object works by specifying a set of differential equations
associated to each synapse (``eqs``) and two rules to specify what should
happen when a presynaptic neuron fires (``pre``) and when a postsynaptic
neuron fires (``post``). The equations should be standard set of equations
in the usual string format. The ``pre`` and ``post`` rules should be a
sequence of statements to be executed triggered on pre- and post-synaptic
spikes. The sequence of statements can be separated by a ``;`` or by
using a multiline string. The reserved symbol ``w`` can be used to refer
to the synaptic weight of the associated synapse.
This framework allows you to implement most STDP rules. Specifying
differential equations and pre- and post-synaptic event code allows for a
much more efficient implementation than specifying, for example, the
spike pair weight modification function, but does unfortunately require
transforming the definition into this form.
There is one restriction on the equations that can be implemented in this
system, they need to be separable into independent pre- and post-synaptic
systems (this is done automatically). In this way, synaptic variables and
updates can be stored per neuron rather than per synapse.
**Example**
::
eqs_stdp = """
dA_pre/dt = -A_pre/tau_pre : 1
dA_post/dt = -A_post/tau_post : 1
"""
stdp = STDP(synapses, eqs=eqs_stdp, pre='A_pre+=delta_A_pre; w+=A_post',
post='A_post+=delta_A_post; w+=A_pre', wmax=gmax)
**STDP variables**
You can access the pre- and post-synaptic variables as follows::
stdp = STDP(...)
print stdp.A_pre
Alternatively, you can access the group of pre/post-synaptic variables
as::
stdp.pre_group
stdp.post_group
These latter attributes can be passed to a :class:`StateMonitor` to
record their activity, for example. However, note that in the case of
STDP acting on a connection with heterogeneous delays, the recent values
of these variables are automatically monitored and these can be
accesses as follows::
stdp.G_pre_monitors['A_pre']
stdp.G_post_monitors['A_post']
**Technical details**
The equations are split into two groups, pre and post. Two groups are created
to carry these variables and to update them (these are implemented as
:class:`NeuronGroup` objects). As well as propagating spikes from the source
and target of ``C`` via ``C``, spikes are also propagated to the respective
groups created. At spike propagation time the weight values are updated.
'''
def __init__(self, C, eqs, pre, post, wmin=0, wmax=Inf, level=0, clock=None, delay_pre=None, delay_post=None):
'''
C: connection object
eqs: differential equations (with units)
pre: Python code for presynaptic spikes
post: Python code for postsynaptic spikes
wmax: maximum weight (default unlimited)
delay_pre: presynaptic delay
delay_post: postsynaptic delay (backward propagating spike)
'''
if get_global_preference('usecstdp') and get_global_preference('useweave'):
from experimental.c_stdp import CSTDP
log_warn('brian.stdp', 'Using experimental C STDP class.')
self.__class__ = CSTDP
CSTDP.__init__(self, C, eqs, pre, post, wmin=wmin, wmax=wmax,
level=level + 1, clock=clock, delay_pre=delay_pre,
delay_post=delay_post)
return
NetworkOperation.__init__(self, lambda:None, clock=clock)
# Convert to equations object
if isinstance(eqs, Equations):
eqs_obj = eqs
else:
eqs_obj = Equations(eqs, level=level + 1)
# handle multi-line pre, post equations and multi-statement equations separated by ;
if '\n' in pre:
pre = flattened_docstring(pre)
elif ';' in pre:
pre = '\n'.join([line.strip() for line in pre.split(';')])
if '\n' in post:
post = flattened_docstring(post)
elif ';' in post:
post = '\n'.join([line.strip() for line in post.split(';')])
# Check units
eqs_obj.compile_functions()
eqs_obj.check_units()
# Get variable names
vars = eqs_obj._diffeq_names
# Find which ones are directly modified (e.g. regular expression matching; careful with comments)
vars_pre = [var for var in vars if var in modified_variables(pre)]
vars_post = [var for var in vars if var in modified_variables(post)]
# additional dependencies are used to ensure that if there are multiple
# pre/post separated equations they are grouped together as one
#
# We should replace with this code here, in case there are no pre/post synaptic variables
#additional_deps =[]
#if len(vars_pre)>0:
# additional_deps.append('__pre_deps='+'+'.join(vars_pre))
#if len(vars_post)>0:
# additional_deps.append('__post_deps='+'+'.join(vars_post))
additional_deps = ['__pre_deps='+'+'.join(vars_pre),
'__post_deps='+'+'.join(vars_post)]
separated_equations = separate_equations(eqs_obj, additional_deps)
if not len(separated_equations) == 2:
raise ValueError('Equations should separate into pre and postsynaptic variables.')
sep_pre, sep_post = separated_equations
for v in vars_pre:
if v in sep_post._diffeq_names:
sep_pre, sep_post = sep_post, sep_pre
break
index_pre = [i for i in range(len(vars)) if vars[i] in vars_pre or vars[i] in sep_pre._diffeq_names]
index_post = [i for i in range(len(vars)) if vars[i] in vars_post or vars[i] in sep_post._diffeq_names]
vars_pre = array(vars)[index_pre]
vars_post = array(vars)[index_post]
# Check pre/post consistency
shared_vars = set(vars_pre).intersection(vars_post)
if shared_vars != set([]):
raise Exception, str(list(shared_vars)) + " are both presynaptic and postsynaptic!"
# Substitute equations/aliases into pre/post code
def substitute_eqs(code):
for name in sep_pre._eq_names[-1::-1]+sep_post._eq_names[-1::-1]: # reverse order, as in equations.py
if name in sep_pre._eq_names:
expr = sep_pre._string[name]
else:
expr = sep_post._string[name]
code = re.sub("\\b" + name + "\\b", '(' + expr + ')', code)
return code
pre = substitute_eqs(pre)
post = substitute_eqs(post)
# Create namespaces for pre and post codes
pre_namespace = namespace(pre, level=level + 1)
post_namespace = namespace(post, level=level + 1)
pre_namespace['clip'] = clip
post_namespace['clip'] = clip
pre_namespace['Inf'] = Inf
post_namespace['Inf'] = Inf
pre_namespace['enumerate'] = enumerate
post_namespace['enumerate'] = enumerate
# freeze pre and post (otherwise units will cause problems)
all_vars = list(vars_pre) + list(vars_post) + ['w']
pre = '\n'.join(freeze(line.strip(), all_vars, pre_namespace) for line in pre.split('\n'))
post = '\n'.join(freeze(line.strip(), all_vars, post_namespace) for line in post.split('\n'))
# Neuron groups
G_pre = NeuronGroup(len(C.source), model=sep_pre, clock=self.clock)
G_post = NeuronGroup(len(C.target), model=sep_post, clock=self.clock)
G_pre._S[:] = 0
G_post._S[:] = 0
self.pre_group = G_pre
self.post_group = G_post
var_group = {} # maps variable name to group
for v in vars_pre:
var_group[v] = G_pre
for v in vars_post:
var_group[v] = G_post
self.var_group = var_group
# Create updaters and monitors
if isinstance(C, DelayConnection):
G_pre_monitors = {} # these get values put in them later
G_post_monitors = {}
max_delay = C._max_delay * C.target.clock.dt
def gencode(incode, vars, other_vars, wreplacement):
num_immediate = num_delayed = 0
reordering_warning = False
incode_lines = [line.strip() for line in incode.split('\n')]
outcode_immediate = 'for _i in spikes:\n'
# delayed variables
outcode_delayed = 'for _j, _i in enumerate(spikes):\n'
for var in other_vars:
outcode_delayed += ' ' + var + '__delayed = ' + var + '__delayed_values_seq[_j]\n'
for line in incode_lines:
if not line.strip(): continue
m = re.search(r'\bw\b\s*[^><=]?=', line) # lines of the form w = ..., w *= ..., etc.
for var in vars:
line = re.sub(r'\b' + var + r'\b', var + '[_i]', line)
for var in other_vars:
line = re.sub(r'\b' + var + r'\b', var + '__delayed', line)
if m:
num_delayed += 1
outcode_delayed += ' ' + line + '\n'
else:
if num_delayed!=0 and not reordering_warning:
log_warn('brian.stdp', 'STDP operations are being re-ordered for delay connection, results may be wrong.')
reordering_warning = True
num_immediate += 1
outcode_immediate += ' ' + line + '\n'
outcode_delayed = re.sub(r'\bw\b', wreplacement, outcode_delayed)
outcode_delayed += '\n %(w)s = clip(%(w)s, %(min)e, %(max)e)' % {'min':wmin, 'max':wmax, 'w':wreplacement}
return (outcode_immediate, outcode_delayed)
pre_immediate, pre_delayed = gencode(pre, vars_pre, vars_post, 'w[_i,:]')
post_immediate, post_delayed = gencode(post, vars_post, vars_pre, 'w[:,_i]')
log_debug('brian.stdp', 'PRE CODE IMMEDIATE:\n'+pre_immediate)
log_debug('brian.stdp', 'PRE CODE DELAYED:\n'+pre_delayed)
log_debug('brian.stdp', 'POST CODE:\n'+post_immediate+post_delayed)
pre_delay_expr = 'max_delay-d'
post_delay_expr = 'd'
pre_code_immediate = compile(pre_immediate, "Presynaptic code immediate", "exec")
pre_code_delayed = compile(pre_delayed, "Presynaptic code delayed", "exec")
post_code = compile(post_immediate + post_delayed, "Postsynaptic code", "exec")
if delay_pre is not None or delay_post is not None:
raise ValueError("Must use delay_pre=delay_post=None for the moment.")
max_delay = C._max_delay * C.target.clock.dt
# Ensure that the source and target neuron spikes are kept for at least the
# DelayConnection's maximum delay
C.source.set_max_delay(max_delay)
C.target.set_max_delay(max_delay)
# create forward and backward Connection objects or SpikeMonitor objects
pre_updater_immediate = STDPUpdater(C.source, C, vars=vars_pre,
code=pre_code_immediate, namespace=pre_namespace, delay=0 * ms)
pre_updater_delayed = DelayedSTDPUpdater(C, reverse=False, delay_expr=pre_delay_expr, max_delay=max_delay,
vars=vars_pre, other_vars=vars_post,
varmon=G_pre_monitors, othervarmon=G_post_monitors,
code=pre_code_delayed, namespace=pre_namespace, delay=max_delay)
post_updater = DelayedSTDPUpdater(C, reverse=True, delay_expr=post_delay_expr, max_delay=max_delay,
vars=vars_post, other_vars=vars_pre,
varmon=G_post_monitors, othervarmon=G_pre_monitors,
code=post_code, namespace=post_namespace, delay=0 * ms)
updaters = [pre_updater_immediate, pre_updater_delayed, post_updater]
self.contained_objects += updaters
vars_pre_ind = dict((var, i) for i, var in enumerate(vars_pre))
vars_post_ind = dict((var, i) for i, var in enumerate(vars_post))
self.G_pre_monitors = G_pre_monitors
self.G_post_monitors = G_post_monitors
self.G_pre_monitors.update(((var, RecentStateMonitor(G_pre, vars_pre_ind[var], duration=(C._max_delay + 1) * C.target.clock.dt, clock=G_pre.clock)) for var in vars_pre))
self.G_post_monitors.update(((var, RecentStateMonitor(G_post, vars_post_ind[var], duration=(C._max_delay + 1) * C.target.clock.dt, clock=G_post.clock)) for var in vars_post))
self.contained_objects += self.G_pre_monitors.values()
self.contained_objects += self.G_post_monitors.values()
else:
# Indent and loop
pre = re.compile('^', re.M).sub(' ', pre)
post = re.compile('^', re.M).sub(' ', post)
pre = 'for _i in spikes:\n' + pre
post = 'for _i in spikes:\n' + post
# Pre code
for var in vars_pre: # presynaptic variables (vectorisation)
pre = re.sub(r'\b' + var + r'\b', var + '[_i]', pre)
pre = re.sub(r'\bw\b', 'w[_i,:]', pre) # synaptic weight
# Post code
for var in vars_post: # postsynaptic variables (vectorisation)
post = re.sub(r'\b' + var + r'\b', var + '[_i]', post)
post = re.sub(r'\bw\b', 'w[:,_i]', post) # synaptic weight
# Bounds: add one line to pre/post code (clip(w,min,max,w))
# or actual code? (rather than compiled string)
if wmax==Inf:
pre += '\n w[_i,:]=clip(w[_i,:],%(min)e,Inf)' % {'min':wmin}
post += '\n w[:,_i]=clip(w[:,_i],%(min)e,Inf)' % {'min':wmin}
else:
pre += '\n w[_i,:]=clip(w[_i,:],%(min)e,%(max)e)' % {'min':wmin, 'max':wmax}
post += '\n w[:,_i]=clip(w[:,_i],%(min)e,%(max)e)' % {'min':wmin, 'max':wmax}
log_debug('brian.stdp', 'PRE CODE:\n'+pre)
log_debug('brian.stdp', 'POST CODE:\n'+post)
# Compile code
pre_code = compile(pre, "Presynaptic code", "exec")
post_code = compile(post, "Postsynaptic code", "exec")
connection_delay = C.delay * C.source.clock.dt
if (delay_pre is None) and (delay_post is None): # same delays as the Connnection C
delay_pre = connection_delay
delay_post = 0 * ms
elif delay_pre is None:
delay_pre = connection_delay - delay_post
if delay_pre < 0 * ms: raise AttributeError, "Presynaptic delay is too large"
elif delay_post is None:
delay_post = connection_delay - delay_pre
if delay_post < 0 * ms: raise AttributeError, "Postsynaptic delay is too large"
# create forward and backward Connection objects or SpikeMonitor objects
pre_updater = STDPUpdater(C.source, C, vars=vars_pre, code=pre_code, namespace=pre_namespace, delay=delay_pre)
post_updater = STDPUpdater(C.target, C, vars=vars_post, code=post_code, namespace=post_namespace, delay=delay_post)
updaters = [pre_updater, post_updater]
self.contained_objects += [pre_updater, post_updater]
# Put variables in namespaces
for i, var in enumerate(vars_pre):
for updater in updaters:
updater._namespace[var] = G_pre._S[i]
for i, var in enumerate(vars_post):
for updater in updaters:
updater._namespace[var] = G_post._S[i]
self.contained_objects += [G_pre, G_post]
def __call__(self):
pass
def __getattr__(self, name):
if name == 'var_group':
# this seems mad - the reason is that getattr is only called if the thing hasn't
# been found using the standard methods of finding attributes, which for var_index
# should have worked, this is important because the next line looks for var_index
# and if we haven't got a var_index we don't want to get stuck in an infinite
# loop
raise AttributeError
if not hasattr(self, 'var_group'):
# only provide lookup of variable names if we have some variable names, i.e.
# if the var_index attribute exists
raise AttributeError
G = self.var_group[name]
return G.state_(name)
def __setattr__(self, name, val):
if not hasattr(self, 'var_group') or name not in self.var_group:
object.__setattr__(self, name, val)
else:
G = self.var_group[name]
G.state_(name)[:] = val
class ExponentialSTDP(STDP):
'''
Exponential STDP.
Initialised with the following arguments:
``taup``, ``taum``, ``Ap``, ``Am``
Synaptic weight change (relative to the maximum weight wmax)::
f(s) = Ap*exp(-s/taup) if s >0
f(s) = Am*exp(s/taum) if s <0
``interactions``
* 'all': contributions from all pre-post pairs are added
* 'nearest': only nearest-neighbour pairs are considered
* 'nearest_pre': nearest presynaptic spike, all postsynaptic spikes
* 'nearest_post': nearest postsynaptic spike, all presynaptic spikes
``wmin=0``
minimum synaptic weight
``wmax``
maximum synaptic weight
``update``
* 'additive': modifications are additive (independent of synaptic weight)
(or "hard bounds")
* 'multiplicative': modifications are multiplicative (proportional to w)
(or "soft bounds")
* 'mixed': depression is multiplicative, potentiation is additive
See documentation for :class:`STDP` for more details.
'''
def __init__(self, C, taup, taum, Ap, Am, interactions='all', wmin=0, wmax=None,
update='additive', delay_pre=None, delay_post=None, clock=None):
if wmax is None:
raise AttributeError, "You must specify the maximum synaptic weight"
wmax = float(wmax) # removes units
eqs = Equations('''
dA_pre/dt=-A_pre/taup : 1
dA_post/dt=-A_post/taum : 1''', taup=taup, taum=taum, wmax=wmax)
if interactions == 'all':
pre = 'A_pre+=Ap'
post = 'A_post+=Am'
elif interactions == 'nearest':
pre = 'A_pre=Ap'
post = 'A_post=Am'
elif interactions == 'nearest_pre':
pre = 'A_pre=Ap'
post = 'A_post=+Am'
elif interactions == 'nearest_post':
pre = 'A_pre+=Ap'
post = 'A_post=Am'
else:
raise AttributeError, "Unknown interaction type " + interactions
if update == 'additive':
Ap *= wmax
Am *= wmax
pre += '\nw+=A_post'
post += '\nw+=A_pre'
elif update == 'multiplicative':
if Am < 0:
pre += '\nw*=(1+A_post)'
else:
pre += '\nw+=(wmax-w)*A_post'
if Ap < 0:
post += '\nw*=(1+A_pre)'
else:
post += '\nw+=(wmax-w)*A_pre'
elif update == 'mixed':
if Am < 0 and Ap > 0:
Ap *= wmax
pre += '\nw*=(1+A_post)'
post += '\nw+=A_pre'
elif Am > 0 and Ap < 0:
Am *= wmax
post += '\nw*=(1+A_pre)'
pre += '\nw+=A_post'
else:
if Am > 0:
raise AttributeError, "There is no depression in STDP rule"
else:
raise AttributeError, "There is no potentiation in STDP rule"
else:
raise AttributeError, "Unknown update type " + update
STDP.__init__(self, C, eqs=eqs, pre=pre, post=post, wmin=wmin, wmax=wmax, delay_pre=delay_pre, delay_post=delay_post, clock=clock)
if __name__ == '__main__':
pass
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