/usr/share/perl5/Bio/DB/GFF/ is in libbio-perl-perl 1.6.923-1.
This file is owned by root:root, with mode 0o755.
..
/usr/share/perl5/Bio/DB/GFF/Adaptor/
/usr/share/perl5/Bio/DB/GFF/Adaptor/ace.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/berkeleydb/
/usr/share/perl5/Bio/DB/GFF/Adaptor/berkeleydb.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/berkeleydb/iterator.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/biofetch.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/biofetch_oracle.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/caching_handle.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/iterator.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/mysql.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/mysqlace.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/mysqlcmap.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/mysqlopt.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/oracle.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/oracleace.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/pg.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/dbi/pg_fts.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/memory/
/usr/share/perl5/Bio/DB/GFF/Adaptor/memory.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/memory/feature_serializer.pm
/usr/share/perl5/Bio/DB/GFF/Adaptor/memory/iterator.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/
/usr/share/perl5/Bio/DB/GFF/Aggregator.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/alignment.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/clone.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/coding.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/gene.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/match.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/none.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/orf.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/processed_transcript.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/so_transcript.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/transcript.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/ucsc_acembly.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/ucsc_ensgene.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/ucsc_genscan.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/ucsc_refgene.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/ucsc_sanger22.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/ucsc_sanger22pseudo.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/ucsc_softberry.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/ucsc_twinscan.pm
/usr/share/perl5/Bio/DB/GFF/Aggregator/ucsc_unigene.pm
/usr/share/perl5/Bio/DB/GFF/Featname.pm
/usr/share/perl5/Bio/DB/GFF/Feature.pm
/usr/share/perl5/Bio/DB/GFF/Homol.pm
/usr/share/perl5/Bio/DB/GFF/RelSegment.pm
/usr/share/perl5/Bio/DB/GFF/Segment.pm
/usr/share/perl5/Bio/DB/GFF/Typename.pm
/usr/share/perl5/Bio/DB/GFF/Util/
/usr/share/perl5/Bio/DB/GFF/Util/Binning.pm
/usr/share/perl5/Bio/DB/GFF/Util/Rearrange.pm