/usr/include/ is in nautilus-actions 3.2.3-1.
This file is owned by root:root, with mode 0o755.
..
/usr/include/nautilus-actions/
/usr/include/nautilus-actions/na-boxed.h
/usr/include/nautilus-actions/na-core-utils.h
/usr/include/nautilus-actions/na-data-types.h
/usr/include/nautilus-actions/na-object.h
/usr/include/nautilus-actions/na-gconf-monitor.h
/usr/include/nautilus-actions/na-object-api.h
/usr/include/nautilus-actions/na-object-item.h
/usr/include/nautilus-actions/na-icontext.h
/usr/include/nautilus-actions/na-ifactory-object-data.h
/usr/include/nautilus-actions/na-iio-provider.h
/usr/include/nautilus-actions/na-object-id.h
/usr/include/nautilus-actions/na-ifactory-object.h
/usr/include/nautilus-actions/na-data-def.h
/usr/include/nautilus-actions/na-ifactory-provider.h
/usr/include/nautilus-actions/na-iexporter.h
/usr/include/nautilus-actions/na-object-menu.h
/usr/include/nautilus-actions/na-data-boxed.h
/usr/include/nautilus-actions/na-dbus.h
/usr/include/nautilus-actions/na-object-action.h
/usr/include/nautilus-actions/na-timeout.h
/usr/include/nautilus-actions/na-iduplicable.h
/usr/include/nautilus-actions/na-extension.h
/usr/include/nautilus-actions/na-gconf-utils.h
/usr/include/nautilus-actions/na-ifactory-provider-provider.h
/usr/include/nautilus-actions/na-object-profile.h
/usr/include/nautilus-actions/na-iimporter.h